- "These howlers make good target practice, eh, Jaden?"
- ―Rosh Penin
Howlers were predators which lived on the moon Yavin 4. They got their name from their method of attack, which was a howl loud enough to stun their prey. Howlers were somewhat reptilian in appearance. They usually crawled on all four limbs, but sometimes when charging, leaping, and clawing, they used two legs for mobility.
- Star Wars: Jedi Knight: Jedi Academy (First appearance)
- Star Wars: Empire at War: Forces of Corruption: Prima Official Game Guide
- The Complete Star Wars Encyclopedia
- Book of Sith: Secrets from the Dark Side