A Shiro's main defense was its spiny shell, which it would withdraw into.
Shiros rolled in mud and collected dirt and seeds between their shell ridges. Tooke-trap plants grew and flourished in those soil pockets, creating the symbiotic "Shiro-trap". The tooke-plants also provided camouflage for Shiros from predators, especially from the saw-toothed grank which was the Shiro's main predator due to its ability to break through tough shells.
Shiros also provided locomotion for the tooke-traps.
- Star Wars Episode I: The Gungan Frontier (First appearance)
- Empire 21: A Little Piece of Home, Part 2