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TR4-121 was a security surveillance camera in the Jedi Temple, specifically the Room of a Thousand Fountains. When Darth Vader, the fallen Jedi Anakin Skywalker, arrived to slaughter the Jedi during Operation: Knightfall in 19 BBY, TR4-121 recorded Skywalker killing Cin Drallig, the Jedi Battlemaster.
When Master Obi-Wan Kenobi and Grand Jedi Master Yoda returned to Coruscant to recalibrate the Jedi beacon, Kenobi saw his former Padawan through the camera. What happened to it afterwards is uncertain.